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Target gene upregulation includes HO-1 (haem oxygenase-1), NQO1 (NAD(P)H quinone oxidoreductase 1), thioredoxin reductase, glutathione peroxidase, and ferritin — a coordinated antioxidant programme that reduces ROS accumulation, lipid peroxidation (MDA reduction approximately 34–44%), and oxidative DNA damage (8-OHdG reduction approximately 28–34%)

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Oxidation is first conducted by DepA (for DON Epimerization), a Type I pyrroloquinoline quinone-dependent alcohol dehydrogenase (PQQ-ADH